A symbiotic origin of magnetoreception in unicellular eukaryotes.
Monteil CL, Vallenet D, Menguy N, Benzerara K, Barbe V, Fouteau S, Cruaud C, Floriani M, Viollier E, Adryanczyk G, Leonhardt N, Faivre D, Pignol D, López-García P, Weld RJ & Lefevre CT
Nature microbiology (2019), 4, 1088-1095

Origin of high Fe contents in modern lacustrine microbialites from Mexico.
Zeyen N, Benzerara K, Menguy N, Brest J, Templeton A, Webb S, Gérard E, Moreira D, Lopez-Garcia P, Tavera R, Morin G.
Geochim Cosmochim Acta
(2019), 253, 201-230

Evidence of high Ca uptake by cyanobacteria forming intracellular CaCO3 and impact on their growth.
De Wever A, Benzerara K, Coutaud M, Caumes G, Poinsot M, Skouri-Panet F, Laurent T, Duprat E, Gugger M
(2019), 17, 676-690.

Exceptional preservation requires fast biodegradation: the case study of thylacocephalan specimens from La Voulte-sur-Rhône (Callovian, Jurassic, France)
Jauvion C, Bernard S, Gueriau P, Mocuta C, Pont S, Benzerara K and Charbonnier S.
(2019), in press.

Pulsed laser-induced heating of mineral phases: implications for the combination/coupling of LIBS and Raman spectroscopy.
Fau A, Beyssac O, Gauthier M, Cousin A, Meslin PY, Benzerara K, Bernard S, Boulliard JC, Maurice S, Gasnault O, Wiens R, Morand M, Rosier P, Garino Y, Pont S   Spectrochimica Acta Part B Atomic Spectroscopy (2019), 160, 105687.

Hyperdiverse archaea near life limits at the polyextreme geothermal Dallol area.
Belilla J, Moreira D, Jardillier L, Reboul G, Benzerara K, López-García JM, Bertolino, López-Archilla AI, López-García P
Nature Ecology & Evolution
(2019), 3, 1552–1561.

Discovery of high abundances of aster-like nanoparticles in pelagic environments: characterization and dynamics.
Colombet J, Billard H, Viguès B, Balor S, Boulé C, Geay L, Benzerara K, Menguy N, Ilango G, Fuster M, Enault F, Bardot C, Gautier V, Sriram Pradeep Ram A, Sime-Ngando T.
Frontiers in Microbiology (2019), 10, 2376.

Sequestration of radionuclides Radium-226 and Strontium-90 by cyanobacteria forming intracellular calcium carbonates.
Mehta N, Benzerara K, Kocar BD, Chapon V.
Environmental Science & Technology (2019), in press

Magnetite Magnetosome Biomineralization in Magnetospirillum magneticum strain AMB-1: a Time Course Study.
Le Nagard L, Zhu X, Yuan H, Benzerara K, Bazylinski D, Fradin C, Swaraj C, Stanescu S, Belkhou R and Hitchcock AP.
Chemical Geology (2019), in press


Active biomineralization of intracellular carbonates by high Ca uptake in cyanobacteria.
Cam N, Benzerara K, Georgelin T, Jaber M, Lambert JF, Poinsot M, Skouri-Panet F, Raimbault E, Cordier L, Jezequel D.
(2018), 16, 49-61

In vitro and in silico evidence of phosphatase diversity in the biomineralizing bacterium Ramlibacter tataouinensis.
Skouri-Panet F, Benzerara K, Cosmidis J, Ferard C, Caumes G, De Luca G, Heulin T, Duprat E.
Frontiers in Microbiology
(2018) 8:2592.

Impact of the cyanobacterium Gloeomargarita lithophora on the geochemical cycles of Sr and Ba
Blondeau M, K. Benzerara, C. Ferard, J-M. Guigner, M. Poinsot, M. Tharaud, L. Cordier, F. Skouri-Panet
Chemical Geology
(2018), 483, 88-97.

Characterization of pustular mats and related Rivularia-rich laminations in oncoids from the Laguna Negra lake (Argentina)
Mlewski E.C., Pisapia C, Gómez F, Lecourt L, Soto Rueda Eliana, Benzerara K, Menez B, Borensztajn S, Jamme F, Refregiers M and Gérard E
Frontiers in Microbiology
(2018) 9: 996.

Amorphous calcium carbonate granules form within an intracellular compartment in calcifying cyanobacteria
Blondeau M, Sachse M Boulogne C Gillet C, Guigner JM, Skouri-Panet F Poinsot M Ferard C, Miot J, Benzerara K
Frontiers in Microbiology

Organic carbon drives transition metal distribution and secondary mineralizations at low temperature in the hydrated mantle-derived oceanic crust.
Ménez B, Pasini V, Guyot F, Benzerara K, Bernard S, Brunelli D


Magnetotactic bacteria are a significant P reservoir in the meromictic Lake Pavin
Rivas-Lamelo S, Benzerara K, Lefèvre CT, Monteil CL, Jézéquel D, Menguy N, Viollier E, Guyot F, Férard C, Poinsot M, Skouri-Panet F, Trcera N, Miot J, Duprat E
Geochemical Perspective Letters
(2017) 5, 35-41.

Single-Cell Resolution of Uncultured Magnetotactic Bacteria via Fluorescence-Coupled Electron Microscopy.
Li JH, Zhang H, Menguy N, Benzerara K, Wang F, Lin X, Chen Z, Pan Y
Applied Environmental Microbiology (2017), 83(12), e00409-17. doi: 10.1128/AEM.00409-17

Description of Gloeomargarita lithophora gen. nov., sp. nov., a thylakoid-bearing basal-branching cyanobacterium with intracellular carbonates, and proposal for Gloeomargaritales ord. nov.
Moreira D; Tavera R; Benzerara K; Skouri-Panet F; Couradeau E; Gérard E; Loussert Fonta E; Novelo E; Zivanovic Y; López-García P.
International Journal of Systematic and Evolutionary Microbiology, In press. doi: 10.1099/ijsem.0.001679

An early-branching freshwater cyanobacterium at the origin of chloroplasts
Ponce-Toledo RI, Deschamps, P, López-García P, Zivanovic Y, Benzerara K, Moreira D.
Current Biology
(2017), 27, 1-6.

Magnetotactic Coccus Strain SHHC-1 Affiliated to Alphaproteobacteria Forms Octahedral Magnetite Magnetosomes
Zhang, H, Menguy, N, Wang, FX, Benzerara, K, Leroy, E, Liu, PY, Liu, WQ, Wang, CL, Pan, YX, Chen, ZB, Li, JH
Frontiers in Microbiology (2017), 8, 969, doi:


Biomineralization patterns of intracellular carbonatogenesis in cyanobacteria: Molecular hypotheses.
Li J, Margaret Oliver I, Cam N, Boudier T, Blondeau M, Leroy E, Cosmidis J, Skouri-Panet F, Guigner JM, Férard C, Poinsot M, Moreira D, Lopez-Garcia P, Cassier-Chauvat C, Chauvat F, Benzerara K
(2016) 6, 10, doi:10.3390/min6010010

Mineralogical diversity in Lake Pavin: connections with water column chemistry and biomineralization processes.
Miot J, Jézéquel D, Benzerara K, Cordier L, Rivas-Lamelo S, Skouri-Panet F, Férard C, Poinsot  M and Duprat E
(2016) 6, 24, doi:10.3390/min6020024

Involvement of microbial mats in early fossilization by decay delay and formation of impressions and replicas of vertebrates and invertebrates
Iniesto M, Buscalioni A, Guerrero MC, Benzerara K, Moreira D, and López-Archilla AI
Scientific reports
(2016) 6: 25716.

Iron mineralogy across the oxycline of a lignite mine lake
Miot J, Lu S, Morin G, Adra A, Benzerara K, Küsel K
Chemical Geology
(2016), 434, 28-42.

Comparative metagenomics unveils functions and genome features of microbialite-associated communities along a depth gradient
Saghaï A, Zivanovic Y, Moreira D, Benzerara K, Bertolino P, Ragon M, Tavera R, López-Archilla AI and López- García P
Environmental Microbiology
(2016), 18, 4990-5004.

Selective uptake of alkaline earth metals by cyanobacteria forming intracellular carbonates
Cam N, Benzerara K, Georgelin T, Jaber M, Lambert JF, Poinsot M, Skouri-Panet F , Cordier L.
Env Sci Technol
(2016) 50, 11654-11662.

Variations in cometary dust composition from Giotto to Rosetta, clues to their formation mechanisms
Engrand C., Duprat J., Dartois E., Benzerara K., Leroux H., Baklouti D., Bardyn A., Briois C., Cottin H., Fischer H., Fray N., Hilchenbach M., Langevin Y., Paquette J., Rynö J., Schulz R., Silén J., Stenzel O., Thirkell L., and the COSIMA team
Monthly Notices of the Royal Astronomical society
(2016) 462, 323-330.


Characterization of Ca-phosphate biomaterials by scanning transmission x-ray microscopy (STXM) at the Ca L2,3-, P L2,3- and C K- edges
Cosmidis J, K. Benzerara, T. Tyliszczak
Acta Biomaterialia
(2015) 12, 260-269.

Formation and transformations of Fe-rich serpentines by aqueous alteration in asteroids: a nanoscale study of the Murray chondrite
Elmaleh A, Bourdelle F, Caste F, Benzerara K, Leroux H, Devouard B
Geochimica et Cosmochimica Acta
(2015) 158, 162-178.

In vitro synthesis of amorphous Mg-, Ca-, Sr- and Ba-carbonates: what do we learn about intracellular calcification by cyanobacteria?
Cam N., T. Georgelin, M. Jaber, J-F. Lambert, K. Benzerara
Geochimica et Cosmochimica Acta 
(2015) 161, 36-49.

Coprolites of Late Triassic carnivorous vertebrates from Poland: an integrative approach.
Zatoń M, Niedźwiedzki G, Marynowski L, Benzerara K, Pott C, Cosmidis J, Krzykawski T and Filipiak P
Palaeogeography, Palaeoclimatology, Palaeoecology
(2015) 430, 21-46.

Emerging frontiers and surprises in Geomicrobiology
A Templeton and K Benzerara
(2015) 11, 423-429.

Metagenome-based diversity analyses suggest a significant contribution of non-cyanobacterial lineages to carbonate precipitation in modern microbialites
A Saghaï, Y Zivanovic, N Zeyen, D Moreira, K Benzerara, P Bertolino, M Ragon, R Tavera, AI López-Archilla and P López-García
Frontiers in Microbiology
(2015) 6:797. doi: 10.3389/fmicb.2015.00797

Preservation in microbial mats: the mineralization through a talc-like mineral of a fish embedded in a microbial sarcophagus
Iniesto, M; Zeyen N; López-Archilla AI; Delgado A.; Guerrero, M.C. and Benzerara K
Frontiers in Earth Science
(2015), doi: 10.3389/feart.2015.00051

Microbial formation of low-T hydrated silicates in modern microbialites from Mexico
Zeyen N, Benzerara K, Li J, Groleau A, Balan E, Robert JL, Tavera R, López-García P, Moreira D
Frontiers in Earth Science
(2015), doi: 10.3389/feart.2015.00064

Calcium-phosphate biomineralization induced by alkaline phosphatase activity in Escherichia coli.
Cosmidis J, Benzerara K, Guyot F, Skouri-Panet F, Férard C, Guigner JM, Babonneau F, Coehlo C
Frontiers in Earth Science
(2015) 3, #00084, DOI=10.3389/feart.2015.00084

Evolution of the macromolecular structure of sporopollenin during thermal degradation as documented by infrared, Raman and C-XANES spectroscopies
S. Bernard, K. Benzerara, O. Beyssac, E. Balan, G.E. Brown Jr
(2015) doi:10.1016/j.heliyon.2015.e00034


Biomineralization of iron-phosphates in the water column of Lake Pavin (Massif Central, France).
Cosmidis J., Benzerara K., Morin G., Busigny V., Lebeau O., Jézéquel D., Noël V., Dublet G., Othmane G
Geochim. Cosmochim. Acta
(2014) 126, 78- 96.

Investigating Microbe-Mineral Interactions: Recent Advances in X-Ray and Electron Microscopy and Redox-Sensitive Methods.
Miot J, Benzerara K., Kappler A.
Annual Review of Earth and Planetary Sciences
(2014) 42: 271-289.

Formation of single domain magnetite by green rust oxidation promoted by microbial anaerobic nitrate-dependent iron oxidation.
Miot J., Li J., Benzerara K, Sougrati M, Ona-Nguema G., Jumas JC, Guyot F
Geochim. Cosmochim. Acta
(2014) 139, 327-343.

Impact of biomineralization on the preservation of microorganisms during fossilization: an experimental perspective
J. Li, S. Bernard, K. Benzerara, T. Allard, J. Cosmidis, J. Moussou, O. Beyssac
Earth Planet. Sci. Lett.
(2014) 400, 113-122.

16 S rDNA-based analysis reveals cosmopolitan occurrence but limited diversity of two cyanobacterial lineages with contrasted patterns of intracellular carbonate mineralization.
Ragon M., Benzerara K., Moreira D., Tavera R. and López-García P.
Frontiers in Microbiology
(2014)  5, #331.

Molecular phylogeny and ultrastructure of Aphelidium aff. Melosirae (Aphelida, Opisthosporidia)
Karpov SA, Mamkaeva MA, Benzerara K, Moreira D, López-García P
(2014) 165, 512-526.

Comparative mineralogy, organic geochemistry and microbial diversity of the Autun black shale and Graissessac coal (France).
Berlendis S., O. Beyssac, S. Derenne, K. Benzerara, C. Anquetil, M. Guillaumet, I. Estève, B. Capelle
International Journal of Coal Geology
(2014) 132, 147-157.

The 2.1 Ga Old Francevillian Biota: Biogenicity, Taphonomy and Biodiversity.
Abderrazak El Albani, Stefan Bengtson, Donald E. Canfield, Armelle Riboulleau, Claire Rollion Bard, Roberto Macchiarelli, Lauriss Ngombi Pemba, Emma Hammarlund, Alain Meunier, Idalina Moubiya Mouele, Karim Benzerara, Sylvain Bernard, Philippe Boulvais, Marc Chaussidon, Christian Cesari, Claude Fontaine, Ernest Chi-Fru, Juan Manuel Garcia Ruiz, François Gauthier-Lafaye, Arnaud Mazurier, Anne Catherine Pierson-Wickmann, Olivier Rouxel, Alain Trentesaux, Marco Vecoli, Gerard J. M. Versteegh, Lee White, Martin Whitehouse, Andrey Bekker
(2014) 9, e99438. doi:10.1371/journal.pone.0099438

Intracellular Ca-carbonate biomineralization is widespread in cyanobacteria

K. Benzerara, F. Skouri-Panet, J. Li, C. Férard, M. Gugger, T. Laurent, E. Couradeau, M. Ragon, J. Cosmidis, N. Menguy, I. Margaret-Oliver, R. Tavera, P. López-García and D. Moreira
Proc Natl. Acad. Sci. USA, (2014) doi:10.1073/pnas.1403510111 link


Nanometer-scale characterization of exceptionally preserved bacterial fossils in Paleocene phosphorites from Ouled Abdoun (Morocco).
Cosmidis, J.; Benzerara, K.; Gheerbrant, E.; et al.
(2013) 11, 139-153.

Multidisciplinary Evidences that Synechocystis PCC6803 Exopolysaccharides Operate in Cell Sedimentation and Protection against Salt  and Metal Stresses.
Jittawuttipoka T., Planchon M., Spalla O.; Benzerara, K, Guyot, F, Cassier-Chauvat C, Chauvat, F
(2013), 8, e55564

Graphite formation by carbonate reduction during subduction.
Galvez M. E., O. Beyssac, I. Martinez, K. Benzerara, C. Chaduteau, B. Malvoisin, J. Malavieille
Nature Geoscience
(2013), 6, 473–477.

Three-dimensional Raman and fluorescence imaging reveal specific carbonate-microbe interactions in modern microbialites.
Gerard E., Menez B., Couradeau E., Moreira D., Benzerara K., Tavera R. and López- García P.
(2013)  doi:10.1038/ismej.2013.81

Sphaerotilus natans, a neutrophilic iron-related sheath- forming bacterium: perspectives for metal remediation strategies.
Seder-Colomina M., G. Morin, K. Benzerara, G. Ona-Nguema, J-J. Pernelle, G. Esposito,
E. van Hullebusch
Geomicrobiology J. 
(2013) doi: 10.1080/01490451.2013.806611

Quantification of the ferric/ferrous iron ratio in silicates by scanning transmission x-ray microscopy at the Fe L2,3 edges
Bourdelle F., K. Benzerara, O. Beyssac, J. Cosmidis, D. Neuville, G.E. Brown Jr, E. Paineau
Contributions to Mineralogy and Petrology 
(2013), 166, 423-434.

Cyanobacterial calcification in modern microbialites at the submicrometer-scale.
Couradeau E., K. Benzerara, E. Gerard, I. Esteve, D. Moreira, R. Tavera, and P. Lopez- Garcia
(2013) 10, 5255-5266.

Low temperature hydrothermal oil and associated biological precursors in serpentinites from Mid-Ocean Ridge.
Pasini V., D. Brunelli, P. Dumas, C. Sandt, J. Frederick, K. Benzerara, S. Bernard, B. Menez
(2013). 10.1016/j.lithos.2013.06.014.
Effect of iron metal and siderite on the durability of simulated archeological glassy material.
Michelin A., Burger E., Leroy E, Foy E, Neff D., Benzerara K., Dillmann P., Gin S.
Corrosion Science
(2013) 76, 403–414.

The link between biomineralization and fossilization of bacteria: Insights from field and experimental studies.
J. Li, K. Benzerara, S. Bernard, O. Beyssac
Chemical Geology 
(2013) 359, 49-69.

Metasomatism and graphite formation at a lithological interface in Malaspina (Alpine Corsica, France).
Galvez, ME.; Martinez, I; Beyssac O; Benzerara K, Agrinier P, Assayag N
Contributions to Mineralogy and Petrology
(2013) 166, 1687-1708.

Microscopy evidence of bacterial microfossils in phosphorite crusts of the Peruvian shelf: implications for phosphogenesis mechanisms

J. Cosmidis, K. Benzerara, N. Menguy, E. Arning
Chemical Geology (2013) 359 10-22


An early-branching microbialite cyanobacterium forms intracellular carbonates.

Couradeau E., Benzerara K., Gérard E., Moreira D., Bernard S., Brown Jr. GE., López- García P.

Science (2012) 336, 459-462.

Micro- and nano-textural evidence of Ti(–Ca–Fe) mobility during fluid–rock interactions in carbonaceous lawsonite-bearing rocks from New Zealand
M.E. Galvez, O. Beyssac, K. Benzerara, N. Menguy, S. Bernard and S. Cox
Contrib Mineral Petrol (2012) 164:895–914

Morphological preservation of carbonaceous plant fossils in blueschist metamorphic rocks from New Zealand
M.E. Galvez, O. Beyssac, K. Benzerara, S. Bernard, N. Menguy, SC Cox, I. Martinez, MR Johnston GE Brown
Geobiology (2012) 10, 118-129

Experimental investigation of the stability of Fe-rich carbonates in the lower mantle
E. Boulard, N. Menguy, AL Auzende, K. Benzerara, H. Bureau, D. Antonangeli, A. Corgne, G. Morard, J. Siebert, JP Perillat, F. Guyot, G. Fiquet
J. Geophysical Research - Solid Earth (2012), 117, B02208

Green rust formation during Fe(II) oxidation by the nitrate-reducing Acidovorax sp. strain BoFeN1.
Pantke C., M. Obst, K. Benzerara, G. Morin, G. Ona-Nguema, U. Dippon, A. Kappler.
Env. Sci. Technol.
(2012) 46, 1439-1446.


Study of the crystallographic architecture of corals at the nanoscale by scanning transmission x-ray microscopy and transmission electron microscopy
K. Benzerara, N. Menguy, M. Obst, J. Stolarski, M. Mazur, T. Tylisczak, G. E. Brown Jr., A. Meibom
Ultramicroscopy, (2011) 111 1268 - 1275 (pdf)

Solenicola setigera is the first characterized member of the abundant and cosmopolitan uncultured marine stramenopile group MAST-3.
Gómez F, Moreira D, Benzerara K. and López-García P.
Environmental Microbiology
(2011) 13, 193-202

A unique skeletal microstructure of the deep-sea micrabaciid scleractinian corals.
Janiszewska K, Stolarski J, Benzerara K, Meibom A, Mazur M, Kitahara M.V. and Cairns S.D.
Journal of Morphology
(2011) 272, 191-203

Significance, mechanisms and environmental implications of microbial biomineralization
Benzerara K., Jennyfer Miot, Guillaume Morin, Feriel Skouri-Panet, Céline Férard
C.R. Geoscience
(2011)  343, 160-167

Study of the crystallographic architecture of corals at the nanoscale by scanning transmission x-ray microscopy and transmission electron microscopy.
Benzerara K., N. Menguy, M. Obst, J. Stolarski, M. Mazur, T. Tylisczak, G. E. Brown Jr., A. Meibom
 (2011) 111,1268-75.

Preservation of protein globules and peptidoglycan in the mineralized cell wall of nitrate-reducing iron(II) oxidizing bacteria: a cryo-electron microscopy study.
Miot J., K. MacLellan, N. Boisset, K. Benzerara
(2011) 9, 459-470.

Influence of uranium on bacterial communities: a comparison of natural uranium-rich soils with controls
Mondani L., K. Benzerara, M. Carrière, R. Christen, Y. Mamindy-Pajany, L. Février, N. Marmier, W. Achouak, P. Nardoux, C. Berthomieu, V. Chapon
PlosOne, 6, e25771.

Biogenic versus metamorphic origins of diverse microtubes in 2.7 Gyrs old volcanic ashes: multi-scale investigations.
 Lepot K., K. Benzerara, P. Philippot
Earth Planet. Sci. Lett., 
(2011) 312, 37-47.

Prokaryotic and eukaryotic community structure in field and cultured microbialites from the alkaline lake Alchichica (Mexico).
Couradeau E., K. Benzerara, D. Moreira, E. Gérard, J. Kazmierczak, R. Tavera & P. López-García
(2011) 6, e28767


XANES, Raman and XRD study of anthracene-based cokes and saccharose- based chars. submitted to high temperature pyrolysis.
Bernard S., O. Beyssac, K. Benzerara, N. Findling, G. Tzvetkov, G.E. Brown Jr.
(2010) 48, 2506-2516.

Multiscale characterization of pyritized plant tissues in blueschist facies metamorphic rocks.
Bernard S., K. Benzerara, O. Beyssac, G.E. Brown Jr.
Geochim. Cosmochim. Acta
(2010) 74, 5054-5068.

Microscopy study of biologically mediated alteration of natural mid-oceanic ridge basalts and magnetic implications
J. Carlut, K. Benzerara, H. Horen, N. Menguy, D. Janots, N. Findling, A. Addad, I. Machouk
J. of Geoph. Res. Biogeosciences (2010), 115, G00G11  


Formation of cell-iron-mineral aggregates by phototrophic and nitrate-dependent anaerobic Fe(II)-oxidizing bacteria.
Schädler S., C. Burkhardt, F. Hegler, K.L. Straub, J. Miot, K. Benzerara, A. Kappler
Geomicrobiology Journal
(2009) 26, 93-103.

Iron biomineralization by neutrophilic iron-oxidizing bacteria.
Miot J., K. Benzerara, G. Morin, A. Kappler, S. Bernard, M. Obst, C. Férard, F. Skouri- Panet, J.M. Guigner, N. Posth, M. Galvez, G.E. Brown Jr, F. Guyot
Geochimica Cosmochimica Acta 
(2009) 73, 696-711.

Transformation of vivianite by anaerobic nitrate-reducing iron-oxidizing bacteria.
Miot J., K. Benzerara, G. Morin, S. Bernard, O. Beyssac, E. Larquet, G. Ona-Nguema, A.   Kappler, F. Guyot
(2009) 7, 373-384.

Looking for traces of life in minerals.
Benzerara K., N. Menguy
CR Palevol
(2009) 8, 617-628.

Extracellular iron biomineralization by photoautotrophic iron-oxidizing bacteria.
 Miot J., K. Benzerara, M. Obst, A. Kappler, F. Hegler, C. Bouchez, F. Guyot, G. Morin.
Applied and Environmental Microbiology
(2009) 75, 5586-5591.

Organic matter heterogeneity in 2.72 Ga stromatolites: alteration versus preservation by sulphur incorporation,
Lepot K., K. Benzerara, G.E. Brown Jr., P. Philippot
Geochimica Cosmochimica Acta
(2009) 73, 6579-6599.

Garnet-filled trails associated with carbonaceous matter mimicking microbial filaments in Archaean basalt.
Lepot K., P. Philippot, K. Benzerara, G.Y Wang
(2009) 7, 1-10.

Ultrastructural and chemical study of modern and fossil sporoderms by Scanning Transmission X-ray Microscopy (STXM).
Bernard S., K. Benzerara, O. Beyssac, G.E. Brown Jr., L. Grauvogel Stamm, P. Duringer
Review of Palaeobotany and Palynology
(2009) 156, 248-261.

Precipitation of amorphous CaCO3 (aragonite-like) and by cyanobacteria: a STXM study of the influence of EPS on the nucleation process.
Obst M., J.J. Dynes, J.R. Lawrence, G.D.W. Swerhone, K. Benzerara, C. Karunakaran, K. Kaznatcheev, T. Tyliszczak, A. P. Hitchcock
Geochimica Cosmochimica Acta
(2009) 73, 4180-4198.

Javaux E.J., K. Benzerara
CR. Palevol  (2009) 8, 605-615


Microbially influenced formation of 2,724-million-year-old stromatolites.
Lepot K., K. Benzerara, G.E. Brown Jr. & P. Philippot
Nature Geoscience
 (2008) 1, 118 - 121.

Nanoscale study of As biomineralization in an acid mine drainage system.
Benzerara K., G. Morin, T.H. Yoon, J. Miot, T. Tyliszczak, C. Casiot, O. Bruneel, F. Farges, and G.E. Brown, Jr.
Geochim. Cosmochim. Acta
(2008) 72,  3949-3963.

Raman mapping using advanced line- scanning systems: geological applications
Bernard S., O. Beyssac, K. Benzerara
Applied Spectroscopy
(2008) 62, 1180-1188.


Exceptional preservation of fossil plant spores in high-pressure metamorphic rocks
S. Bernard, K. Benzerara, O. Beyssac, N. Menguy, F. Guyot, G. E. Brown Jr. and B. Goffé
Earth and Planetary Science Letters, (2007), 262, 257-272 (pdf)
Highligt Nature Geoscience, 18 Oct 2007

Alteration of submarine basaltic glass from the Ontong Java Plateau: A STXM and TEM study
K. Benzerara, N. Menguy, N.R. Banerjee, T. Tyliszczak, G.E. Brown Jr. and F. Guyot
Earth and Planetary Science Letters (2007), 260, 187-200 (pdf)

Creating habitable zones, at all scales, from planets to mud micro- habitats, on earth and on mars. S
Nisbet E., Zahnle K, Gerasimov MV, Helbert J, Jaumann R, Hofmann BA, Benzerara K, Westall F.
(2007) 129 (1-3): 79-121.

Water, life, and planetary geodynamical evolution.
van Thienen P, Benzerara K, Breuer D, Gillmann C, Labrosse S, Lognonne P, Spohn T
(2007) 129 (1-3): 167-203.


icrobial diversity on the Tatahouine meteorite
Benzerara K., V. Chapon, D. Moreira, P. López-García, T. Heulin, F. Guyot. M
Meteoritics and Planetary Sciences(2006) 41, 1249-1265.

Nanometer-Scale Chemical Heterogeneities of Black Carbon Materials and Their Impacts on PCB Sorption Properties: Soft X-ray Spectromicroscopy Study.
Yoon T.H., Benzerara K., Ahn S., Luthy R.G., Tyliszczak T. and Brown, G.E. Jr.
Env. Sci.
&Technol. (2006) 40, 5923-5929.

Search for microbial signatures within human and microbial calcifications using soft X-ray spectromicroscopy.
 Benzerara K., V.M. Miller, G. Barell, V. Kumar, J. Miot, G.E. Brown, Jr., and J.C. Lieske
Journal of Investigative Medicine
(2006)   54, 367-379.

The desert of Tataouine: an extreme environment that hosts a wide diversity of microorganisms and radiotolerant bacteria.
Chanal A., Chapon V., Benzerara K., Barakat M., Christen R., Chevenet F., Achouak W., Barras F., and Heulin T.
Environ. Microbiol.
 (2006).  8 (3): 514-525.

Soft X-ray Microscopy and Spectroscopy at the Molecular Environmental Science Beamline at the Advanced Light Source.
Bluhm H., Andersson K., Araki T., Benzerara K., Brown G.E., Dynes J.J., Ghosal S., Gilles M.K., Hansen H.-C., Hemminger J.C., Hitchcock A.P., Ketteler G., Kilcoyne A.L.D., Kneedler E., Lawrence J.R., Leppard G.G., Majzlam J., Mun B.S., Myneni S.C.B., Nilsson A., Ogasawara H., Ogletree D.F., Pecher K., Salmeron M., Shuh D.K., Tonner B., Tyliszczak T., Warwick T., Yoon T.H.
J. Elec.Related Phenom. (2006) 150, Spectros. and 86-104.

Nanoscale detection of organic signatures in carbonate microbialites
K. Benzerara, N. Menguy, P. Lopez-Garcıa, T.H. Yoon , J. Kazmierczak,
T. Tyliszczak, F. Guyot, and G.E. Brown, Jr.
Proc Natl. Acad. Sci. USA, (2006), 103, 9440-9445 (pdf)


Bacterial diversity and carbonate precipitation in the giant microbialites from the highly alkaline Lake Van, Turkey.
P., Kazmierczak J., Benzerara K., Kempe S., Guyot F., Moreira D.
9, 263-274.

Nanoscale environments associated with bioweathering of a Mg-Fe-pyroxene
K. Benzerara, T.H. Yoon, N. Menguy, T. Tyliszczak, and G.E. Brown, Jr.
Proc Natl. Acad. Sci. USA, (2005), 102, 979-982 (pdf)

TEM study of a silicate-carbonate-microbe interface prepared by focused ion beam milling
K. Benzerara, N. Menguy, F. Guyot, Ch. Vanni and Ph. Gillet
Geochimica et Cosmochimica Acta, (2005), 69, 1413-1422 (pdf)


Scanning Transmission X-ray Microscopy Study of Microbial Calcification
Benzerara K., Yoon T.-H., Tyliszczak T., Constantz B., Spormann A.M., and Brown, G.E. Jr.
(2004) 2, ,249-259.

In Situ Characterization of Aluminum-Containing Nanoparticle- Aqueous Suspensions Using Scanning Transmission X-ray Microscopy.
Yoon T.-H., Johnson S.B., Benzerara K., Doyle C.S., Tyliszczak T., Shuh D.K., and Brown G.E., Jr.
(2004) 20, 10361-10366.

Experimental colonization and alteration of orthopyroxene by the pleomorphic bacteria Ramlibacter tataouinensis,
K. Benzerara, M. Barakat, N. Menguy, F. Guyot, G. De Luca, C. Audrain, T. Heulin
Geomicrobiology Journal, (2004), 21, 341 – 349 (pdf)

Biologically controlled precipitation of calcium phosphate by Ramlibacter tataouinensis
K. Benzerara, N. Menguy, F. Guyot, F. Skouri, G. de Luca, M. Barakat and T. Heulin
Earth and Planetary Science Letters, (2004), 228, 439-449 (pdf)


Nannobacteria like calcite single crystals at the surface of the Tatahouine meteorite,
Benzerara K, Menguy, N, Guyot F, Dominici C, Gillet P,
Proc Natl. Acad. Sci. USA, (2003), 100, 7438-7442 (pdf)


Cristobalite inclusions in the Tatahouine achondrite: Implications for the shock conditions.
Benzerara K., J.A. Barrat, F. Guyot, P. Gillet and M. Lesourd.
 American Mineralogist
(2002) 87, 1250-1256.

Aqueous alteration in the North West Africa 817 (NWA 817) Martian Meteorite
Gillet, Ph. Barrat, J.A, Deloule, E., Wadhwa, M., Jambon, A., Sautter, V., Crozaz, G., Devouard, B., Neuville, D., Benzerara, K., Lesourd, M.
Earth and  Planetary Science Letters
(2002) 203, 431-444.


Description of new shock-induced phases in the meteorites of Shergotty, Zagami, Nakhla and Chassigny.
Malavergne, V., Guyot, F., Benzerara K., and Martinez, I.
(2001) 36, 1297-1306.


Bacteria in the Tatahouine meteorite: nanometric-scale life in rocks.
Gillet, P., Barrat, J.A., Heulin, T., Achouak, W., Lesourd, M., Guyot, F., and Benzerara K.
Earth and Planetary Science Letters (2000) , 175, 161-167.


Karim BENZERARA     Modifié le 15 Septembre 2018